This chapter connects the major transitions to evolvability and its evolution. It reviews a three-pulse model of the conditions that make the evolution of complexity possible. It specifically outlines the life up to the evolution of fully equipped prokaryote cells, the phase of microbial evolution, and the third phase that sees the evolution of complex development. It compares the prokaryote and multicellular evolvability. The deeply obscure problem of precellular evolution is also evaluated. This chapter shows that individual properties of developmental mechanisms are significant to evolvability, but they are not all that is important.
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